Genus Pulmonaria L. - Lungworts
L., Sp. PI., ed. 1 (1753) 153; Gen. PI., ed. 5 (1754) 65.
Lungworts are perennial herbs that form clumps or rosettes. They are covered in hairs of varied length and stiffness, and sometimes also bear glands. The underground parts consist of a slowly creeping rhizome with adventitious roots. Flowering stems are unbranched, rough, covered with bristly hairs, usually not exceeding 25–30cm, with a few exceptions (P. mollis, P. vallarsae). The stems are usually upright, or slightly spreading.
The leaves are arranged in rosettes. The blades are usually large, from narrowly lanceolate to oval, with the base ranging from heart shaped to very gradually narrowing, and can have a sharply pointed or blunt tip. Leaf margin is always entire, but in some species and forms can be rather wavy. Basal leaves are carried on stalks that can be short or longer than the leaf blade in various species. Stem leaves are smaller and often narrower, and are unstalked or clasping the stem. All leaves are covered with hairs that are usually bristly, or occasionally soft. The leaves are often prominently spotted in black and blue, or sometimes in pale green, or unspotted.
The inflorescence is a terminal scorpioid cyme, with bracts. Lungwort flowers are heterostylous, with two distinct forms of flower within each species; those with short stamens and long styles ("pin" flowers) and those with long stamens and short styles ("thrum" flowers), with the former usually being larger and more showy. The calyx is hairy, 5-lobed, tubular or funnel-shaped, enlarging as the fruit ripens. The corolla is funnel-shaped and consists of a long, cylindrical tube and a limb with five shallow lobes. Within the corolla throat, five tufts of hairs alternate with the stamens to form a ring. The colour of corolla varies from purple, violet or blue to shades of pink and red, or sometimes white. The colour of the flower in bud is often pink, which then changes as the flower matures. The stamens and style are included within the corolla and not protruding.
The nutlets are smooth, egg-shaped, brownish, up to 4.5 mm long and 3 mm wide, each containing a single seed. Up to four nutlets per flower are produced, ripening mostly in summer.
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Pulmonaria officinalis L.